Μελέτη της ανωμειοκαινικής απολιθωματοφόρου θέσης σπονδυλωτών της Κρυοπηγής και άλλων θέσεων της Κασσάνδρας Χαλκιδικής. Συστηματική, ταφονομία, παλαιοοικολογία, βιοχρονολόγηση
Περίληψη
In the present dissertation thesis the fossil vertebrate material from the Kryopigi and other localities of the Kassandra Peninsula of Chalkidiki, N. Greece, have been studied. The total number of studied specimens exceeds 8000 and each one has been collected during the systematic excavation research in the area the past 15 years. This study includes the systematic paleontology, the taphonomical analysis of the material, the paleoecological analysis and the biochronology of the various faunas. These localities have been found in terrestrial deposits of Antonios Formation and Triglia Formation, both dated to Late Miocene. The Kryopigi (KRY), the Kassandra-Chomateri (KCH) and the Polychrono (KPO) localities belong to Triglia Formation. Specimens that come from the Kassandra-Kryopigi (KKR), the “sand deposits of Kassandra” (FRK, KSN, AMPVG), the Aghia Paraskevi (PAR) and Chelona-Siviri (KSC) have been found in deposits of the Antonios Formation. Furthermore, in the “mad deposits of Kassndra” plant remains have been found. The material from Kassandra Peninsula belongs to 46 different taxa. The collection of the Kryopigi locality includes 32 taxa of mammals, reptiles and birds. The fauna from the Kassandra-Chomateri includes 5 taxa. The fauna of the “sand deposits of Kassandra” 9 taxa have been identified and in the Polychrono locality only one species. Four new taxa are proposed in the Kryopigi: the hipparion H. phlegrae the pig Propotamochoerus aegaeus sp. nov. the bird Otis hellenica and the bovid Prostrepsiceros axiosi minus subsp. nov. The fauna from the “sand deposits of Kassndra” is considered of Vallesian age. The species Tetralophodon longirostris is referred for the first time and the genus Urmiatherium has never been referred earlier from the Balkans apart from the Turolian Samos faunas. The Kryopigi fauna is dated to the second half of the Turolian, in late MN12 or early MN13. The various species appear to be more evolved than those from the faunas of Ravin des Zouaves-5 and Prochoma in the Axios Valley. Probably the Kryopigi fauna is more evolved from the Perivolaki fauna and more primitive than that of Dytiko localities. Αn age between 7.3-6.4 Ma is suggested for the Kryopigi fauna. The fauna from the Kassandra-Chomateri is dated to Turolian and maybe earlier than that of Kryopigi. The Polychrono locality is also dated to Turolian. At least 268 individuals have been estimated in the Kryopigi fauna. The bones in the Kryopigi locality accumulated due to carnivore activity during repeating dry periods and buried fast and close to their original place of death during floods. Paleoecological analysis includes the estimation of habitat score on the bovid and hipparion metapodials, the guild structure of the carnivores, the cenogram analysis of the Kryopigi fauna, and the estimation of the faunal similarity. All the results show more or less the same picture about the Kryopigi paleoecology. The environment was open, savanna-like with hot and dry climate. The presence of Varanus sp. in the fauna is indicative of an annual mean temperature 15ο C. Some animals in the Kryopigi fauna were well adapted in more forested environment.
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Abel, O. 1904. Über einen Fund von Sivatherium giganteum bei Adrianopel. [s.n.]
(Kaiserlich-königlichen Hof-und Staatsdruckerei).
Abel, O. 1926. Die Geschichte der Equiden auf dem Boden Noramerikas. Verhandlungen
Zoologisch-Botanischen Gesellschaft, 74:159-164.
Ahmed, Z. 2003a. A new species of the genus Propotamochoerus from the lower Chinji
formation of lower Siwalik, District Chakwal, Punjab, Pakistan. Biologia (Pakistan),
(1 & 2):71-76.
Ahmed, Z. 2003b. A new suid species Propotamochoerus khani from Siwaliks of Pakistan.
Biologia (Pakistan), 49(1 & 2):9-20.
Ahmed, Z. 2008. The occurence of the genus Propotamochoerus in Dhokpathan formation of
the Middle Siwaliks of Pakistan. Biologia (Pakistan), 54(1):17-32.
Alba, D. M., E. Delson, G. Carnevale, S. Colombero, M. Delfino, P. Giuntelli, M. Pavia, and
G. Pavia. 2014. First joint record of Mesopithecus and cf. Macaca in the Miocene of
Europe. J Hum Evol, 67:1-18.
Alcalá, L., and C. M. Escorza. 1998. Modelling diagenetic bone fractures. Bulletin de la
Société Géologique de France, 169(1):101-108.
Alexeiev, A. 1930. Die obersarmatische Fauna von Eldar. I. Achtiaria borissiaki nov. sp.
Trav. du Musee geol. Acad. Sc. URSS, 8:167-204.
Alexejev, A. 1915. Animaux fossiles du village Novo-Elizavetovka. Tipografiya "Technik",
Odessa, 453 p.
Anderson, K., and L. Werdelin. 2005. Carnivora from the late Miocene of Lantian, China.
Vertebrate PalAsiatica, 43(4):256-271.
Andrée, J. 1921. Rhinocerotiden aus dem Unterpliocän von Samos. Paläontologische
Zeitschrift, 3(1):189-212.
Andrée, J. 1926. Neue Cavicornier aus dem Pliocän von Samos. Palaeontographica,
(6):135-175.
Andrews, P., T. Harrison, E. Delson, R. L. Bernor, and L. Martin. 1996. Distribution and
biochronology of European and Southwest Asian Miocene catarrhines. The evolution
of Western Eurasian Neogene mammal faunas:168-207.
Arambourg, C., and J. Piveteau. 1929. Les Vertébrés du Pontien de Salonique. Annales de
Paléontologie, 18:59-138.
Athanassiou, A. 1996. Contribution to the study of the fossil mammals of Thessaly. PhD.
Thesis, University of Athens [in Greek], 1-350 p.
Barone, R. 1976. Anatomie Comparée des Mammifères Domestiques. Tome I: Ostéologie.
Vigot Fréres, Paris, 428 p.
Bernor, R. L. 1985. Systematic and evolutionary relationships of the hipparionine horses from
Maragheh, Iran (Late Miocene, Turolian age). Laboratoire de Paléontologie de
l'Université des Sciences et Techniques du Languedoc.
Bernor, R. L., M. Armour-Chelu, T. M. Kaiser, and R. S. Scott. 2003. An Evaluation of the
Late MN 9 (Late Miocene, Vallesian Age), Hipparion Assemblage from Rudabánya
(Hungary): Systematic Background, Functional Anatomy and Paleoecology.
Coloquios de Paleontología, Vol. Ext. 1:35-45.
Bernor, R. L., T. M. Kaiser, S. V. Nelson, and L. Rook. 2011. Systematics and Paleobiology
of Hippotherium malpassii n. sp. (Equidae, Mammalia) from the latest Miocene of
Baccinello V3 (Tuscany, Italy). Bollettino della Società Paleontologica Italiana,
(3):175-208.
Bernor, R. L., N. Solounias, C. Swisher, and A. Van Couvering. 1996. The Correlation of
Three Classical “Pikermian” Mammal Faunas—Maragheh, Samos, and Pikermi—with
the European MN Unit System, p. 137-154. In R. L. Bernor, V. Fahlbush, and H.-W.
Mittman (eds.), The evolution of Western Eurasian Neogene mammal faunas.
Columbia University Press, New York.
Bernor, R. L., and H. Tobien. 1989. Two small species of Cremohipparion (Equidae,
Mamm.) from Samos, Greece Mitt. Bayer. Staatsslg. Paläont. Hist. Geol., 29:207-226.
Bibi, F., and E. S. Güleç. 2008. Bovidae (Mammalia: Artiodactyla) from the Late Miocene of
Sivas, Turkey. Journal of Vertebrate Paleontology, 28(2):501-519.
Boev, Z., and G. Koufos. 2000. Presence of Pavo bravardi (Gervais, 1849)(Aves,
Phasianidae) in the Ruscinian locality of Megalo Emvolon, Macedonia, Greece.
Geologica Balcanica, 30(1/2):69-69.
Boev, Z., and G. D. Koufos. 2006. The late Miocene vertebrate locality of Perivolaki,
Thessaly, Greece. 2. Aves, p. 11-22. In G. D. Koufos (ed.), The late Miocene
vertebrate locality of Perivolaki, Thessaly, Greece.Volume 276. Palaeontographica
Boev, Z., G. Lazaridis, and E. Tsoukala. 2013. Otis hellenica sp. nov., a new Turolian bustard
(Aves: Otididae) from Kryopigi (Chalkidiki, Greece). Geologica Balcanica, 42(1-
:59-65.
Bohlin, B. 1926. Die familie Giraffidae mit besonderer berücksichtigung der fossilen formen
aus China. Paleontologia Sinica. Geological Survey of China, 178 p.
Bohlin, B. 1935a. Cavicornier der hipparion-fauna nord-Chinas. Paleontologia Sinica, C,
(4):1-166.
Bohlin, B. 1935b. Tsaidamotherium hedini, n.g., n. sp. Ein Einhörniger Ovibovine, aus den
Tertiären Ablagerungen aus der Gegend des Tossun nor, Tsaidam. Geografiska
Annaler:66-74.
Böhme, M. 2003. The Miocene Climatic Optimum: evidence from ectothermic vertebrates of
Central Europe. Palaeogeography, Palaeoclimatology, Palaeoecology, 195(3-4):389-
Böhme, M., A. Ossig, and H. Küchenhoff. 2006. New method to estimate paleoprecipitation
using fossil amphibians and reptiles and the Middle and Late Miocene precipitation
gradients in Europe. Geophys J R Astron Soc, 34(6):425-428.
Böhme, M., and D. Vasilyan. 2014. Ectothermic vertebrates from the late Middle Miocene of
Gratkorn (Austria, Styria). Palaeobiodiversity and Palaeoenvironments, 94(1):21-40.
Boon-Kristkoiz, E., and A. Kristkoiz. 1999. Order Lagomorpha, p. 259-262. In G. E. Rössner
and K. Heissig (eds.), The Miocene Land Mammals of Europe. Verlag Dr. Friedrich
Pfeil, Munchen.
Borissiak, A. 1915. Mammifères fossiles de Sébastopol. Memmoires du Comite Geologique,
:105-154.
Bosscha-Erdbrink, D. P. 1977. Distribution in time and space of three giraffid genera with
Turolian representatives at maragheh in NW Iran. PROCEEDINGS OF THE
KONINKLIJKE NEDERLANDSE AKADEMIE VAN WETENSCHAPPEN SERIES
B-PALAEONTOLOGY GEOLOGY PHYSICS CHEMISTRY ANTHROPOLOGY,
(5):337-355.
Bouvrain, G. 1978. Protragelaphus theodori n. sp. (Mammalia, Artiodactyla, Bovidae) du
Miocène de Macédoine (Grèce). Géologie Méditerranéenne, 5:229-236.
Bouvrain, G. 1979. Un genre nouveau de Bovidè de la fin du Miocène. Bulletin de la Société
Géologique de France, 21(4):507-511.
Bouvrain, G. 1980. Le genre Palaeoreas (Mammalia, Artiodactyla, Bovidae), systématique et
extension géographique. Paläontologische Zeitschrift, 54(1-2):55-65.
Bouvrain, G. 1982. Révision du genre Prostrepsiceros Major 1891 (Mammalia, Bovidae).
Paläontologische Zeitschrift, 56(1-2):113-124.
Bouvrain, G. 1988. Les Tragoportax (Bovidae, Mammalia) des gisements du Miocène
supérieur de Dytiko (Macédoine, Grèce). Annales de Paléontologie, 74:43-63.
Bouvrain, G. 1994. Un bovidé du Turolien inférieur d’Europe orientale: Tragoportax
rugosifrons. Annales de Paléontologie, 80(1):61-87.
Bouvrain, G. 1996. Les gazelles du Miocène supérieur de Macédoine, Grèce. NEUES
JAHRBUCH FUR GEOLOGIE UND PALAONTOLOGIE ABHANDLUNGEN,
(1):111-132.
Bouvrain, G. 1997. Les Bovidés du Miocène supérieur de Pentalophos (Macédoine, Grèce).
Münchner Geowissenschaftliche abhandlungen, Reihe A, Geologie und Paläontologie,
:5-22.
Bouvrain, G. 2001. Les bovides (Mammalia, Artiodactyla) des gisements du Miocene
superieur de Vathylakkos (Grece du Nord). Neues Jahrbuch für Geologie und
Paläontologie - Abhandlungen, 220(2):225-244.
Bouvrain, G., and L. de Bonis. 1984. Le genre Mesembriacerus (Bovidae, Artiodactyla,
Mammalia): un oviboviné primitif du Vallésien (Miocène supérieur) de Macédoine
(Grèce). Palaeovertebrata, 14(4):201-223.
Bouvrain, G., and L. de Bonis. Ruminants (Mammalia, Artiodactyla: Tragulidae, Cervidae,
Bovidae) des gisements du Miocène supérieur (Turolien) de Dytiko (Grèce). Annales
de Paléontologie, 93:121-147.
Bouvrain, G., S. Sen, and H. Thomas. 1995. Parurmiatherium rugosifrons Sickenberg, 1932,
un Ovibovinae (Bovidae) du Miocène supérieur d'Injana (Djebel Hamrin, Irak).
Geobios, 28(6):719-726.
Bullet, P. 1942. Beiträge zur kenntnis des Gebisses von Varanus salvator Laur. Zoologischen
Museum der Universität Zürich:139-192.
Cannon, M. D. 2013. NISP, Bone Fragmentation, and the Measurement of Taxonomic
Abundance. Journal of Archaeological Method and Theory, 20(3):397-419.
Chang, Y.-p. 1974. Miocene suids from Kaiyuan, Yunnan and Linchu, Shantung. Vertebrata
Palasiatica, 12(2):117-123.
Christaras, V. 1984. Geology of the bauxitic ores of the Mt. Katsika (Chalkidiki), PhD Thesis,
Scientific Annals, School of Geology, University of Thessaloniki, 7: 23, 209 pp.(in
Greek).
Ciric, A. 1957. Pikermiska fauna iz okoline Titovog Velesa. Bulletin du Muséum d'Histoire
naturelle du Pays serbe A, 8:1-82.
Clavel, J., G. Merceron, L. Hristova, N. Spassov, D. Kovachev, and G. Escarguel. 2012. On
Mesopithecus habitat: Insights from late Miocene fossil vertebrate localities of
Bulgaria. J Hum Evol, 63(1):162-179.
Clos, L. M. 1995. A New Species of Varanus (Reptilia: Sauria) from the Miocene of Kenya.
Journal of Vertebrate Paleontology, 15(2):254-267.
Colbert, E. H. 1935. Distributional and Phylogenetic Studies on Indian Fossil Mammals: The
Phylogeny of the Indian Suidae and the Origin of the Hippopotamidae. American
Museum of Natural History, 799.
Conrad, J. L., A. M. Balcarcel, and C. M. Mehling. 2012. Earliest example of a giant monitor
lizard (Varanus, Varanidae, Squamata). PLoS One, 7(8):e41767.
Cooke, H. B. S. 1982. A preliminary appraisal of fossil Suidae from Sahabi, Libya. Garyounis
Scientific Bulletin:71-82.
Cope, E. D. 1868. On the origin of genera. Proceedings of the Academy of Natural Sciences
of Philadelphia, 20:242-300.
Costeur, L. 2005. Cenogram analysis of the Rudabánya mammalian community:
palaeoenvironmental interpretations. Palaeontographia Italica, 90:303-307.
Dayan, T., and D. Simberloff. 1996. Patterns of size separation in carnivore communities.
Carnivore behavior, ecology, and evolution, 2:243-266.
Dayan, T., E. Tchernov, Y. Yom-Tov, and D. Simberloff. 1989. Ecological character
displacement in Saharo-Arabian Vulpes: outfoxing Bergmann's rule. Oikos:263-272.
de Beaumont, G. 1986. Les Carnivores (Mammifères) du Néogène de Höwenegg/Hegau,
Baden-Württemberg. Carolinea, 44:35-45.
de Beaumont, G., and P. Mein. 1972. Recherches sur le genre Plioviverrops Kretzoi
(Carnivora,? Hyaenidae). Comptes Rendus des Séances, SPHN Genéve, NS,
(25):383-394.
de Bonis, L. 2004. Carnivores hyaenidés du Miocène supérieur de Turquie. Zona
arqueológica(4):108-117.
de Bonis, L. 2005. Carnivora (Mammalia) from the late Miocene of Akkaşdagi, Turkey.
Geodiversitas, 27(4):567-589.
de Bonis, L., and G. Bouvrain. 1996. Suidae du Miocène supérieur de Grèce. Bulletin du
Muséum national d'histoire naturelle. Section C, Sciences de la terre, paléontologie,
géologie, minéralogie, 18(1):107-132.
de Bonis, L., G. Bouvrain, D. Geraads, and G. Koufos. 1990. New remains of Mesopithecus
(Primates, Cercopithecoidea) from the late Miocene of Macedonia (Greece), with the
description of a new species. Journal of Vertebrate Paleontology, 10(4):473-483.
de Bonis, L., G. Bouvrain, D. Geraads, and G. D. Koufos. 1992. A skull of Hystrix primigenia
from the late Miocene of Macedonia (Greece). Neues Jahrbuch für Geologie und
Paläontologie, Monatshefte, 1992(2):75-87.
de Bonis, L., G. Bouvrain, D. Geraads, and G. D. Koufos. 1997. New material of
Mesopithecus (Mammalia, Ceropithecidae) from the late Miocene of Macedonia,
Greece. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen, 5:255-266.
de Bonis, L., G. Bouvrain, G. Koufos, and P. Tassy. 1995. Un crâne de chalicothère
(Mammalia, Perissodactyla) du Miocène supérieur de Macédoine (Grèce): remarques
sur la phylogénie des Chalicotheriinae. Palaeovertebrata, 24(1-2):135-176.
de Bonis, L., and G. Koufos. 1981. A new hyaenid (Carnivora, Mammalia) from the Vallesian
(late Miocene) of northern Greece. Scientific Annals, Faculty Physics and
Mathematics, Univ. Thessaloniki, 21:79-94.
de Bonis, L., and G. D. Koufos. 1991. The late Miocene small carnivores of the lower Axios
valley (Macedonia, Greece). Geobios, 24:361-379.
de Bonis, L., S. Peigné, A. Likius, H. T. Mackaye, P. Vignaud, and M. Brunet. 2005.
Hyaenictitherium minimum, a new ictithere (Mammalia, Carnivora, Hyaenidae) from
the Late Miocene of Toros-Menalla, Chad. Comptes Rendus Palevol, 4(8):671-679.
de Mecquenem, R. 1924. Contribution à l'étude des fossiles de Maragha. Annales de
Paléontologie, 13:135-160.
de Mecquenem, R. 1925. Contribution à l'étude des fossiles de Maragha. Annales de
Paléontologie, 14:1-34.
deBraga, M. 1990. Anatomical and functional changes between terrestrial varanoid lizards
and aquatic mosasaurs, McGill University, Montreal 197 p.
Delfino, M., D. M. Alba, R. Carmona, A. Lujan, and J. Robles. 2011. European monitor
lizards (Anguimorpha, Varanidae, Varanus): new materials and new perspectives.
Journal of Vertebrate Paleontology, 31:97-98.
Delfino, M., D. M. Alba, A. H. Luján, R. Carmona, J. Abella, J. M. Robles, J. Galindo, and S.
Almécija. 2012. Varanus marathonensis Weithofer, 1888: morphology, systematics,
and paleobiogeography of the European monitor lizards, 4 th Interntional Geologica
Belgica Meeting, Moving Plates and Melting Icecaps. Processes and Forcing Factors
in Geology, 156.
Delson, E. 1973. Fossil colobine monkeys of the circum-Mediterranean region: and the
evolutionary history of the Cercopithecidae (Primates, Mammalia), Columbia
University New York.
Delson, E. 1994. Evolutionary history of the colobine monkeys in paleoenvironmental
perspective. Colobine Monkeys: Their Ecology, Behaviour and Evolution, Cambridge
University Press, Cambridge:11-43.
Delson, E., C. J. Terranova, W. L. Jungers, E. J. Sargis, N. G. Jablonski, and P. C. Dechow.
Body mass in Cercopithecidae (Primates, Mammalia): estimation and scaling in
extinct and extant taxa. Anthropological Papers of the American Museum of Natural
History(83):1-159.
Eisenmann, V. 1995. What metapodial morphometry has to say about some Miocene
Hipparions, p. 148-163. In E. S. Vrba, G. H. Denton, T. C. Partridge, and L. H.
Burckle (eds.), Paleoclimate and Evolution, with Emphasis on Human Origins. Yale
University Press, New Haven and London.
Eisenmann, V. 1998. Folivores et tondeurs d’herbe: forme de la symphyse mandibulaire des
equidés et des tapiridés (Perissodactyla, Mammalia). Geobios, 31:113-123.
Eisenmann, V., M. T. Alberdi, C. d. Giuli, and U. Staesche. 1988. Methodology, p. 1-71. In
M. O. Woodburne and P. Y. Sondaar (eds.), Studying Fossil Horses: Collected Papers
After the New York International Hipparion Conference, 1981.Volume 1. E. J. Brill
Archive, New York.
Eisenmann, V., and P. Sondaar. 1998. Pliocene vertebrate locality of Çalta, Ankara, Turkey.
Hipparion. Geodiversitas, 20(3):409-439.
Estes, R., K. de Queiroz, and J. Gauthier. 1988. Phylogenetic relationships within Squamata,
p. 119-281. In R. Estes and G. K. Pregill (eds.), Phylogenetic relationships of the
lizard families. Stanford University Press.
Evans, A. R., M. Fortelius, J. Jernvall, and J. T. Eronen. Dental ecomorphology of extant
European carnivorans. Current Trends in Dental Morphology Research: 13th
International Symposium on Dental Morphology. University of Lodz Press, Lodz:223-
Flynn, J. J., and H. Galiano. 1982. Phylogeny of early Tertiary Carnivora, with a description
of a new species of Protictis from the middle Eocene of northwestern Wyoming,
Forsyth-Major, C. 1888. Sur un gisement d'ossements fossiles dans l'île de Samos
contemporains de l'âge de Pikermi. Comptes Rendus de l’Académie des Sciences,
:1178-1182.
Forsyth-Major, C. 1903. New Carnivora from the middle Miocene of La Grive St-Alban.
Geological Magazine, ns, 1:534-538.
Fortelius, M. [coordinator] 2014. New and Old Worlds Database of Fossil Mammals (NOW).
University of Helsinki.
Fortelius, M., J. van der Made, and R. L. Bernor. 1996. Middle and Late Miocene Suoidea of
central Europe and the eastern Mediterranean: Evolution, biogeography, and
paleoecology, p. 348-377. In R. L. Bernor, V. Fahlbush, and H.-W. Mittmann (eds.),
The evolution of Western Eurasian Neogene mammal faunas. Columbian University
Press, New York.
Gallai, G., and L. Rook. 2011. Propotamochoerus provincialis (Gervais, 1859)(Suidae,
Mammalia) from the latest Miocene (late Messinian; MN13) of Monticino Quarry
(Brisighella, Emilia-Romagna, Italy). Bollettino della Società Paleontologica Italiana,
(1):29-34.
Galloway, A. 1999. The biomechanics of fracture production, p. 35-62. In A. Galloway (ed.),
Broken Bones: Anthropological Analysis of Blunt Force Trauma. Charles C Thomas,
Springfield, Illinois.
Garevski, R. 1976. Weiterer Beitrag zur Kenntnis der Pikermifauna Mazedoniens. Der
Mastodonschädel von der Umgebung des Dorfes Dolni Disan (Negotino). Posebno
Izdanie Musei Macedonici Scientiarum Naturalium, 7:21-26.
Garevski, R. 1997. Ein Schädel samt Unterkiefer von Choerolophodon pentelici Gaudry &
Lartet aus der Pikermifauna von Makedonien. Geološki Zbornik, 12:247-264.
Gasparik,M. 2004. Proboscidean remains from the Pannonian of Rudabánya. Palaeontographia Italica, 90:181-192.
Gaudry, A. 1862–1867. Animaux fossiles et géologie de l’Attique. Paris, Savy, 475.
Gaudry, A. 1873. Animaux fossiles du Mont Luberon (Vaucluse). F. Savy, Paris:112.
Gaudry, A., and E. Lartet. 1856. Résultats des recherches paléontologiques entreprises dans
l’Attique sous les auspices de l’Académie. Comptes-rendus hebdomadaires des
séances de l'Académie des Sciences, 43:271-274.
Gaziry, A. W. 1976. Jungtertiäre Mastodonten aus Anatolien (Türkei). Geologisches
Jahrbuch, 22:3-143.
Gentry, A. W. 1971. The earliest goats and other antelopes from the Samos Hipparion fauna.
Bulletin of the British Museum (Natural History) Geology, 20(6):1-296.
Geraads, D. 1974. Les Giraffidés du Miocène supérieur de la région de Thessalonique
(Grèce), Universite de Paris VI, Paris, 1–102 p.
Geraads, D. 1978. Les palaeotraginae (giraffidae, mammalia) du miocene superieur de la
region de Thessalonique (Grece). Géologie Méditerranéenne, 5:269-276.
Geraads, D. 1994. Les gisements de mammifères du Miocène supérieur de Kemiklitepe,
Turquie: 8. Giraffidae. Bull Mus Nat Hist Nat, Paris C, 16:159-172.
Geraads, D. 2009. Giraffidae (Mammalia) de la fin du Néogène de la République de
Macédoine (ARYM). Geodiversitas, 31(4):893-908.
Geraads, D. 2013. Large Mammals from the late Miocene of Çorakyerler, Çankırı, Turkey.
Acta zoologica bulgarica, 65(3):381-390.
Geraads, D., and E. Güleç. 1999a. A Bramatherium skull (Giraffidae, Mammalia) from the
Late Miocene of Kavakdere (Central Turkey). Biogeographic and phylogenetic
implications. Mineral Res. Expl. Bul, 121:51-56.
Geraads, D., and E. Güleç. 1999b. On some spiral-horned antelopes (Mammalia:
Artiodactyla: Bovidae) from the Late Miocene of Turkey, with remarks on their
distribution. Paläontologische Zeitschrift, 73(3-4):403-409.
Geraads, D., T. Kaya, and S. Mayda. 2005a. Late Miocene large mammals from Yulafl,
Thrace region, Turkey, and their biogeographic implications. Acta Palaeontologica
Polonica, 50(3):523.
Geraads, D., and G. D. Koufos. 1990. Upper Miocene Rhinocerotidae (Mammalia) from
Pentalophos-1, Macedonia, Greece. Palaeontographica Abteilung A, 210:151-168.
Geraads, D., and N. Spassov. 2008. A new species of Criotherium (Bovidae, Mammalia)
from the late Miocene of Bulgaria. Hellenic Journal of Geosciences, 43:21-27.
Geraads, D., N. Spassov, and R. Garevski. 2008. New specimens of Propotamochoerus
(Suidae, Mammalia) from the late Miocene of the Balkans. Neues Jahrbuch für
Geologie und Paläontologie-Abhandlungen, 248(1):103-113.
Geraads, D., N. Spassov, L. Hristova, G. N. Markov, and T. Tzankov. 2011. Upper Miocene
mammals from Strumyani, South-Western Bulgaria. Geodiversitas, 33(3):451-484.
Geraads, D., N. Spassov, and D. Kovachev. 2001. New Chalicotheriidae (Perissodactyla,
Mammalia) from the Late Miocene of Bulgaria. Journal of Vertebrate Paleontology,
(3):596-606.
Geraads, D., N. Spassov, and D. Kovachev. 2003. Palaeoreas lindermayeri (Wagner,
(Mammalia, Bovidae) from the upper Miocene of Bulgaria, and a revision of the
species. Geodiversitas, 25(2):405-415.
Geraads, D., N. Spassov, and D. Kovachev. 2005b. Giraffidae (Artiodactyla, Mammalia)
from the Late Miocene of Kalimantsi and Hadjidimovo, Southwestern Bulgaria.
Geologica Balcanica, 35(1-2):11-18.
Geraads, D., E. Tsoukala, and N. Spassov. 2007. A skull of Ancylotherium (Chalicotheriidae,
Mammalia) from the late Miocene of Thermopigi (Serres, N. Greece) and the
relationships of the genus. Journal of Vertebrate Paleontology, 27(2):461-466.
Gervais, P. 1849. Zoologie et Paléontologie Française. Paris: Bertrand.
Gervais, P. 1859. Nouvelles recherches sur les animaux vertebres dont ou trouve les
ossements enfouis dans le sol de la France. Zoologie et Paléontologie Françaises, 8:1-
Giaourtsakis, I. X. 2003. Late Neogene Rhinocerotidae of Greece: distribution, diversity and
stratigraphical range. Deinsea, 10:235-253.
Giaourtsakis, I. X. 2009. The Late Miocene mammal faunas of the Mytilinii Basin, Samos
Island, Greece: New collection. 9. Rhinocerotidae. Beitrage zur Paläontologie,
:157-187.
Giaourtsakis, I. X., and G. D. Koufos. 2009. The late Miocene mammal faunas of the Mytilini
Basin, Samos Island, Greece: new collection. 10. Chalicotheriidae. Beiträge zur
Paläontologie Wien, 31:189-205.
Giaourtsakis, I. X., G. Theodorou, S. Roussiakis, A. Athanassiou, and G. Iliopoulos. 2006.
Late Miocene horned rhinoceroses (Rhinocerotinae, Mammlia) from Kerassia
(Euboea, Greece). Neues Jahrbuch für Geologie und Paläontologie-Abhandlungen,
(3):367-398.
Ginsburg, L. 1988. Contribution à l’étude du gisement Miocène supérieur de Montredon
(Hérault). 4: les artiodactyles Suidae. Palaeovertebrata, Mémoire extraordinaire,
:57-64.
Godina, A. Y. 1975. On new finds of Sivatheriinae on the territory of the USSR. Journal of
the Palaeontological Society of India, 20:21-25.
Godina, A. Y. 2002. On the taxonomy and evolution of Samotherium (Giraffidae,
Artiodactyla). Paleontological Journal, 36(4):395-402.
Göhlich, U. B. 1998. Elephantoidea (Proboscidea, Mammalia) aus dem Mittel- und
Obermiozän der Oberen Süβwassermolasse Süddeutschlands: Odontologie und
Osteologie. Münchner Geowissenschaftliche abhandlungen, Reihe A, Geologie und
Paläontologie, 36:1-245.
Göhlich, U. B., and K. Huttunen. 2009. The early Vallesian vertebrates of Atzelsdorf (Late
Miocene, Austria) 12. Proboscidea. Annalen des Naturhistorischen Museums in Wien
A, 111:635-646.
Golpe-Posse, J. M. 1980. Notas sobre la dentición anterior de Microstonys antiquus (Kaup,
. Acta Geologica Hispanica, 15(4):109-111.
Gray, J. E. 1821. On the natural arrangement of vertebrose animals. The London Medical
Repository Monthly Journal and Review, 15:296-310.
Gregory, W. K. 1912. Notes on the principles of quadrupedal locomotion and the mechanism
of the limbs in hoofed animals. Annals of the New York Academy of Sciences,
:267-294.
Gromova, V. 1949. Istoriya loshadei (roda Equus) v starom svete; chast II, Evolyutsiya i
klassifikatsiya roda. Trudy Paleontologicheskogo Instituta, 17:1-163.
Gromova, V. 1952. Le genre Hipparion. Bureau de Recherches géologiques et Minières,
C.E.D.P., 12:1-288.
Guy, M., J. Bornovas, J.-J. Bizon, and G. Bizon. 1969. Geological map of Greece 1:50 000,
Peninsula of Kassandra. Institut de Geologie et de Recherche du Sous-sol, Athenes.
Hünermann, K. A. 1968. Die Suidae (Mamamlia, Artiodactyla) aus den Dinotheriensanden
(Unterpliozän. Birkhäuser.
Hamilton, W. R. 1978. Fossil giraffes from the Miocene of Africa and a revision of the
phylogeny of the Giraffoidea. Philosophical Transactions of the Royal Society of
London. Series B, Biological Sciences, 283:165-229.
Hammer, Ø., and D. A. Harper. 2006. Paleontological data analysis. Blackwell Publishing,
p.
Hammer, Ø., D. A. Harper, and P. Ryan. 2001. PASΤ: Paleontological Statistics Software
Package for Education and Data Analysis Palaeontologia Electronica, 4(1):1-9.
Harrison, T., and E. Delson. Mesopithecus sivalensis from the Late Miocene of the Siwaliks.
American journal of physical anthropology:126.
Hayek, L.-A. C., R. L. Bernor, N. Solounias, and P. Steigerwald. 1991. Preliminary studies of
hipparionine horse diet as measured by tooth microwear. Annales zoologici
Fennici:187-200.
Hensel, R. 1862. Über die Reste einiger Säugetierarten von Pikermi in der Münchener
Sammlung. Monatsberichte der Akademie der Wissenschaften, 27:560-569.
Hertler, C., and R. Volmer. 2008. Assessing prey competition in fossil carnivore
communities—a scenario for prey competition and its evolutionary consequences for
tigers in Pleistocene Java. Palaeogeography, Palaeoclimatology, Palaeoecology,
(1):67-80.
Hooijer, D. A. 1954. Pleistocene vertebrates from Celebes. VIII. Dentition and skeleton of
Celebochoerus heekereni Hooijer. Zoologische Verhandelingen, 24(1):1-46.
Howell, F. C., and G. Petter. 1985. Comparative observations on some middle and upper
Miocene hyaenids, Genera: Percrocuta Kretzoi, Allohyaena Kretzoi, Adcrocuta
Kretzoi (Mammalia, Carnivora, Hyaenidae). Geobios, 18(4):419-491.
Hristova, L. 2009. Ontogeny and variability in the cheek region of hipparions from the Late
Miocene locality Hadzhidimovo-1, Southwest Bulgaria. Rivista Italiana di
Paleontologia e Stratigrafia, 115(1):125-132.
Hristova, L., D. Geraads, G. N. Markov, and N. Spassov. 2013. Late Miocene Mammals from
Kocherinovo, Southwestern Bulgaria. Acta zoologica bulgarica, 65(4):517-529.
Hristova, L., D. Kovachev, and N. Spassov. 2002. The Hipparions (Equidae, Mammalia) from
the Upper Miocene locality Hadzhidimovo, SW Bwlgaria. Rev. Bulg. GeoI. Society,
(1-3):89-98.
Hristova, L., D. Kovachev, and N. Spassov. 2003. Hipparion brachypus Hensel, 1862 from
Hadjidimovo, southwestern Bulgaria (Late Miocene). Comptes rendus de l' Academie
bulgare des Sciences, 56(2):77-84.
I.C.V.G.A.N. 2005. Nomina Anatomica Veterinaria (fifth edition). Hannover, Columbia,
Gent, Sapporo, International Committee on Veterinary Gross Anatomical Nomenclature:165.
Iliopoulos, G. 2003. The Giraffidae (Mammalia, Artiodactyla) and the study of the history
and chemistry of fossil mammal bone from the late Miocene of Kerassia (Euboea
Island, Greece). PhD Thesis, University of Leicester, UK, 144 p.
Jablonski, N. G., D. Su, J. Kelly, L. J. Flynn, and X. Ji. 2011. The Mio-Pliocene colobine
monkey, Mesopithecus, in China. American journal of physical anthropology,
:174-174.
Jafarzadeh, R., D. S. Kostopoulos, and J. Daneshian. 2012. Skull reconstruction and ecology
of Urmiatherium polaki (Bovidae, Mammalia) from the upper Miocene deposits of
Maragheh, Iran. Paläontologische Zeitschrift, 86(1):103-111.
Küllmer, O., and C. Doukas. 1995. The vertebrate locality of Maramena (Macedonia, Greece)
at the Turolian-Ruscinian Boundary (Neogene). 6-The deciduous dentition of
Mesopithecus pentelicus Wagner (Primates, Mammalia). Münchner Geowiss. Abh,
:65-74.
Kaya, T. T., S. Mayda, D. S. Kostopoulos, M. C. Alcicek, G. Merceron, A. Tan, S.
Karakütük, A. K. Giesler, and R. S. Scott. 2012. Şerefköy-2, a new late Miocene
mammal locality from the Yatağan Formation, Muğla, SW Turkey. Comptes Rendus
Palevol, 11(1):5-12.
Köhler, M. 1987. Boviden des türkischen Miozäns (Känozoikum und Bräunkohlen der
Türkei). Paleontologia i Evolució, 21:133-246.
Koken, E. 1885. Ueber fossile Saugetiere aus China. Palaeontologische Abhandlungen, 2:31-
Koliadimou, K., and G. D. Koufos. 1991. The Hystricidae from the Pleistocene of Macedonia
(Greece) and a review of the European representatives of the family. Bulletin
Geologial Society of Greece, 25(2):453-471.
Kondopoulou, D., L. de Bonis, G. D. Koufos, and S. Sen. 1993. Paleomagnetic data and
biostratigraphy of the Middle Miocene vertebrate locality of Thymiana, Chios island,
Greece, p. 676-687, Proceedings of the 2nd Congress of the Hellenic Geophysical
Union, Florina.
Kondopoulou, D., S. Sen, G. D. Koufos, and L. de Bonis. 1990. Magneto-and biostratigraphy
of the late Miocene mammalian locality of Prochoma (Macedonia, Greece). Paleont. i
Evolucio, 24(25):135-139.
Konidaris, G. E. 2013. Palaeontological and biostratigraphical study of the Neogene
Proboscidea from Greece. PhD thesis, Aristotle University of Thessaloniki,
Thessaloniki, Greece, 326 p.
Konidaris, G. E., and G. D. Koufos. 2009. The Late Miocene Mammal Faunas of the Mytilinii
Basin, Samos Island, Greece: New Collection. 8. Proboscidea. Beitr. Paläeont, 31:139-
Konidaris, G. E., and G. D. Koufos. 2013. Late Miocene Proboscidea (Mammalia) from
Macedonia and Samos Island, Greece: preliminary results. Paläontologische
Zeitschrift, 87(1):121-140.
Konidaris, G. E., G. D. Koufos, D. S. Kostopoulos, and G. Merceron. 2014a. Taxonomy,
biostratigraphy and palaeoecology of Choerolophodon (Proboscidea, Mammalia) in
the Miocene of SE Europe-SW Asia: implications for phylogeny and biogeography.
Journal of Systematic Palaeontology.
Konidaris, G. E., S. Roussiakis, G. Theodorou, and G. D. Koufos. 2014b. The Eurasian
occurrence of the shovel-tusker Konobelodon (Mammalia, Proboscidea) as illuminated
by its presence in the late Miocene of Pikermi (Greece). Journal of Vertebrate
Paleontology, 34(6):1437-1453.
Korotkevich, E. L. 1978. Samotherium maeoticum new species from the meotian of the
northern black sea area. Vestnik Zoologii, 4:9-18.
Kostopoulos, D. S. 2000. Functional morphology and palaeoecological adaptations of
Nisidorcas planicornis (Bovidae, Mammalia) from the late Miocene. Münchner
geowissenschaftliche Abhandlungen, 39:33-44.
Kostopoulos, D. S. 2004. Revision of some late Miocene spiral horned antelopes (Bovidae,
Mammalia). Neues Jahrbuch für Geologie und Paläontologie-Abhandlungen,
(2):167-190.
Kostopoulos, D. S. 2005. The Bovidae (Mammalia, Artiodactyla) from the late Miocene of
Akkaşdagi, Turkey. Geodiversitas, 27(4):747-791.
Kostopoulos, D. S. 2006. The late Miocene vertebrate locality of Perivolaki, Thessaly,
Greece. 9. Cervidae and Bovidae. Palaeontographica Abteilung A, 276(1-6):151-183.
Kostopoulos, D. S. 2009a. Contribution to the systematics and phylogeny of Prostrepsiceros
vallesiensis Bouvrain, 1982 (Mammalia, Bovidae). Geodiversitas, 31(4):879-891.
Kostopoulos, D. S. 2009b. The Late Miocene mammal faunas of the Mytilinii Basin, Samos
Island, Greece: new collection. 13. Giraffidae. Beitr. Paläeont, 31:299-343.
Kostopoulos, D. S. 2009c. The Late Miocene Mammal faunas of the Mytilinii basin, Samos
Island, Greece: new collection. 14. Bovidae. Beitrage zur Paläontologie, 31:345-389.
Kostopoulos, D. S. 2009d. The Pikermian Event: Temporal and spatial resolution of the
Turolian large mammal fauna in SE Europe. Palaeogeography, Palaeoclimatology,
Palaeoecology, 274(1):82-95.
Kostopoulos, D. S. 2014. Taxonomic re-assessment and phylogenetic relationships of
Miocene homonymously spiral-horned antelopes. Acta Palaeontologica Polonica.
Kostopoulos, D. S. in press. Palaeontology of the Late Miocene Mammal Localities of Nikiti
(Chalkidiki Peninsula, Macedonia, Greece). The Nikiti-2 Artiodactyla revised.
Geobios.
Kostopoulos, D. S., and R. L. Bernor. 2011. The Maragheh bovids (Mammalia, Artiodactyla):
systematic revision and biostratigraphic-zoogeographic interpretation. Geodiversitas,
(4):649-708.
Kostopoulos, D. S., and S. Karakütük. 2015. Late Miocene bovids from Şerefköy-2, SW
Turkey, and their position within the sub-Paratethyan biogeographic province. Acta
palaeontologia Polonica, 60(1):49-66.
Kostopoulos, D. S., K. K. Koliadimou, and G. D. Koufos. 1996. The giraffids (Mammalia,
Artiodactyla) from the late Miocene mammalian localities of Nikiti (Macedonia,
Greece). Palaeontographica Abteilung A, 239(1-3):61-88.
Kostopoulos, D. S., and G. D. Koufos. 1996. Late Miocene bovids (Mammalia, Artiodactyla)
from the locality “Nikiti-1”(NKT), Macedonia, Greece. Annales de Paléontologie,
(4):251-300.
Kostopoulos, D. S., and G. D. Koufos. 1999. The Bovidae (mammalia, artiodactyla) of the
“Nikiti-2” (NIK) faunal assemblage (Chalkidiki Peninsula, N. Greece). Annales de
Paléontologie, 85(3):193-218.
Kostopoulos, D. S., and G. D. Koufos. 2006. The late Miocene vertebrate locality of
Perivolaki, Thessaly, Greece. 8. Giraffidae. Palaeontographica Abteilung A, 276(1-
:135-149.
Kostopoulos, D. S., and G. Saraç. 2005. Giraffidae (Mammalia, Artiodactyla) from the late
Miocene of Akkaşdağı, Turkey. Geodiversitas, 27(4):735-745.
Kostopoulos, D. S., S. Sen, and G. D. Koufos. 2003. Magnetostratigraphy and revised
chronology of the late Miocene mammal localities of Samos, Greece. International
Journal of Earth Sciences, 92(5):779-794.
Kostopoulos, D. S., N. Spassov, and D. Kovachev. 2001. Contribution to the study of
Microstonyx: evidence from Bulgaria and the SE European populations.
Geodiversitas, 23(3):411-437.
Koufos, G. D. 1977. New findings of Mastodonts from Macedonia. Scientific Annals, Faculty
of Physics and Mathematics, University of Thessaloniki, 17:97-115.
Koufos, G. D. 1987a. Chasmaporthetes bonisi, a new hyenid (Carnivora, Mammalia) from
the late Miocene of Macedonia (Greece). Bulletin de la Société Géologique de France,
(5):913-920.
Koufos, G. D. 1987b. Study of the Pikermi hipparions. I: generalities and taxonomy. Bulletin
du Muséum national d'histoire naturelle. Section C, Sciences de la terre, paléontologie,
géologie, minéralogie, 9(2):197-252.
Koufos, G. D. 1987c. Study of the Pikermi Hipparions. II: Comparisons and odontograns.
Bulletin du Muséum national d'histoire naturelle. Section C, Sciences de la terre,
paléontologie, géologie, minéralogie, 9(3):327-363.
Koufos, G. D. 1987d. Study of the Turolian Hipparions of the lower Axios Valley
(Macedonia, Greece): 1. Locality Ravin des Zouaves-5 (RZO). Geobios, 20(3):293-
Koufos, G. D. 1987e. Study of turolian hipparions of the lower Axios valley (Macedonia,
Greece) 2. Locality Prochoma-1 (PXM). Paläontologische Zeitschrift, 61(3-4):339-
Koufos, G. D. 1988. Study of the Turolian Hipparions of the Lower Axios Valley
(Macedonia, Greece): 4. Localities of Dytiko Paleovertebrata, 18:187-239.
Koufos, G. D. 2000. Revision of the late Miocene carnivores from the Axios valley,
Macedonia, Greece. Münchner geowissenschaftliche Abhandlungen, 39:51-92.
Koufos, G. D. 2006a. The large mammals from the Miocene/Pliocene locality of Silata,
Macedonia, Greece with implications about the latest Miocene palaeoecology.
Beitrage zur Paläontologie, 30:293-313.
Koufos, G. D. 2006b. The late Miocene mammal locality of Perivolaki, Thessaly, Greece. 3.
Primates. Palaeontographica Abt A, 276:23-37.
Koufos, G. D. 2006c. The late Miocene vertebrate locality of Perivolaki, Thessaly, Greece. 4.
Carnivora. Palaeontographica Abteilung A, 276(1-6):39-74.
Koufos, G. D. 2006d. The late Miocene vertebrate locality of Perivolaki, Thessaly, Greece. 5.
Proboscidea, Rhinocerotidae. Palaeontographica Abteilung A, 276:75-80.
Koufos, G. D. 2006e. The Neogene mammal localities of Greece: Faunas, chronology and
biostratigraphy. Hellenic Journal of Geosciences, 41(1):183-214.
Koufos, G. D. 2006f. Palaeoecology and chronology of the Vallesian (late Miocene) in the
Eastern Mediterranean region. Palaeogeography, Palaeoclimatology, Palaeoecology,
(2):127-145.
Koufos, G. D. 2008a. Carnivores from the early/middle Miocene locality of Antonios
(Chalkidiki, Macedonia, Greece). Geobios, 41(3):365-380.
Koufos, G. D. 2008b. The Miocene cercopithecid primates of Greece. Giornate di
Paleontologia, Edizione VIII:129-131.
Koufos, G. D. 2009a. The genus Mesopithecus (Primates, Cercopithecidae) in the late
Miocene of Greece. Bolletino della Società Paleontologica Italiana.
Koufos, G. D. 2009b. The Late Miocene Mammal Faunas of the Mytilinii Basin, Samos
Island, Greece: New Collection. 5. Carnivora. Beiträge zur Paléontologie, 31:57-105.
Koufos, G. D. 2009c. The Neogene cercopithecids (Mammalia, Primates) of Greece.
Geodiversitas, 31(4):817-850.
Koufos, G. D. 2011. The Miocene carnivore assemblage of Greece. Estudios Geologicos,
(2):291-320.
Koufos, G. D. 2012a. New material of Carnivora (Mammalia) from the Late Miocene of
Axios Valley, Macedonia, Greece. Comptes Rendus Palevol, 11(1):49-64.
Koufos, G. D. 2012b. New material of Chalicotheriidae (Perissodactyla, Mammalia) from the
Late Miocene of Axios Valley, Macedonia (Greece) with the description of a new
species. Annales de Paléontologie, 98(3):203-224.
Koufos, G. D., L. de Bonis, D. S. Kostopoulos, L. Viriot, and T. D. Vlachou. 2004.
Mesopithecus (Primates, Cercopithecidae) from the Turolian locality of Vathylakkos 2
(Macedonia, Greece). Paläontologische Zeitschrift, 78(1):213-228.
Koufos, G. D., and K. Koliadimou. 1993. Two lagomorphs from the Pliocene of Macedonia,
Greece. Bulletin of the Geological Society of Greece, 28:7-29.
Koufos, G. D., and G. E. Konidaris. 2011. Late Miocene carnivores of the Greco-Iranian
Province: Composition, guild structure and palaeoecology. Palaeogeography,
Palaeoclimatology, Palaeoecology, 305(1):215-226.
Koufos, G. D., D. S. Kostopoulos, and G. Merceron. 2009a. The late Miocene mammal
faunas of the Mytilinii Basin, Samos Island, Greece: New Collection. 17.
Palaeoecology–Palaeobiogeography. Beitrage zur Paläontologie, 31:409-430.
Koufos, G. D., D. S. Kostopoulos, and I. A. Sylvestrou. 2006a. The late Miocene vertebrate
locality of Perivolaki, Thessaly, Greece. 10. Chronology. Palaeontographica Abteilung
A:185-200.
Koufos, G. D., D. S. Kostopoulos, and T. D. Vlachou. 2009b. The Late Miocene Mammal
Faunas of the Mytilinii Basin, Samos Island, Greece: New Collection. 16.
Biochronology. Beiträge zur Paléontologie, 31:397–408.
Koufos, G. D., D. S. Kostopoulos, T. D. Vlachou, and G. E. Konidaris. 2011. A synopsis of
the late Miocene mammal fauna of Samos Island, Aegean Sea, Greece. Geobios,
(2):237-251.
Koufos, G. D., and J. K. Melentis. 1984. The late Miocene (Turolian) mammalian fauna of
Samos island (Greece). Study of the collection of the Palaeontological Museum of
Mytilinii, Samos. 2. Equidae. Scientific Annals of the Faculty of Physics and
Mathematics of the University of Thessaloniki,, 24:47-78.
Koufos, G. D., G. Merceron, D. S. Kostopoulos, and I. A. Sylvestrou. 2006b. The late
Miocene vertebrate locality of Perivolaki, Thessaly, Greece. 11. Palaeoecology and
Palaeobiogeography. Palaeontographica Abteilung A, 276:201-221.
Koufos, G. D., N. Spassov, and D. Kovatchev. 2003. Study of Mesopithecus (Primates,
Cercopithecidae) from the late Miocene of Bulgaria. Palaeontographica Abteilung
A:39-91.
Koufos, G. D., and G. E. Syrides. 1997. A new Early/Middle Miocene mammal locality from
Macedonia, Greece. Comptes Rendus de l'Académie des Sciences-Series IIA-Earth
and Planetary Science, 325(7):511-516.
Koufos, G. D., G. E. Syrides, and K. K. Koliadimou. 1991. A pliocene primate from
Macedonia (Greece). J Hum Evol, 21(4):283-294.
Koufos, G. D., and T. D. Vlachou. 2005. Equidae (Mammalia, Perissodactyla) from the late
Miocene of Akkaşdagi, Turkey. Geodiversitas, 27(4):633-705.
Kovachev, D. 2012a. A complete skeleton of Adcrocuta eximia (Roth and Wagner, 1854)
from the Upper Maeotian (Turolian) of Hadzhidimovo, SW Bulgaria. Geologica
Balcanica, 41(1-3):77-95.
Kovachev, D. 2012b. A porcupine skeleton of Hystrix (Hystrix) primigenia (Wagner) from
the Upper Maeotian (Turolian) of Hadzhidimovo, SW Bulgaria Geologica Balcanica,
(1-3):3-20.
Kovachev, D., and N. Atanasova. 2008. On the presence of genus Hystrix (Rodentia) in the
Upper Maeotian from the region south of Hadzhidimovo town, Blagoevgrad district.
Rev. Bulg. GeoI. Society, 69(1-3):21-26.
Kretzoi, M. Materialen zur phylogenetischen klassifikation der Aeluroideen. 10th
International Zoological Congress, Budapest, 1927:1293-1355.
Kretzoi, M. 1938. Die Raubtiere von Gombaszög nebst einer Ubersicht der Gesamtfauna (Ein
Beitrag zur Stratigraphie des Altquartaers). Annales Musei Nationalis Hungarici, 31.
Kretzoi, M. 1951. The Hipparion fauna from Csákvár. Foldtani Közlony, 81:438-441.
Kretzoi, M. 1952. Die Raubtiere der Hipparionfauna von Polgárdi. Annales Instituti Geologici
Publici Hungarici, 40(3):1-41.
Kurtén, B. 1982. Status of the fossil hyaenids Ιctitherium viverrinum and Τhalassictis robusta
(Mammalia). ZEITSCHRIFT FUR GEOLOGISCHE WISSENSCHAFTEN, 10(7):1009-1018.
Lacombat, F. 2006. Pleistocene Rhinoceroses in Mediterranean Europe and in Massif Central
(France). COURIER-FORSCHUNGSINSTITUT SENCKENBERG, 256:57-69.
Latreille, P. A. 1800. Histoire Naturelle des Salamandres de France, Précédéed’un Tableau
Methodique des autres Reptiles Indigénes, Villier, Paris, 61 p.
Lazaridis, G., and E. Tsoukala. 2014a. A Case of Abnormal Deciduous Tooth Morphology of
a Late Miocene Hipparionine Horse. Acta zoologica bulgarica, 66(2):293-296.
Lazaridis, G., and E. Tsoukala. 2014b. Hipparion phlegrae, sp. nov. (Mammalia,
Perissodactyla): a new species from the Turolian locality of Kryopigi (Kassandra,
Chalkidiki, Greece). Journal of Vertebrate Paleontology, 34(1):164-178.
Lazaridis, G., and E. Tsoukala. 2014c. Tetralophodon longirostris (Kaup, 1832) from Late
Miocene of the Kassandra peninsula (Chalkidiki, Greece). Scientific Annals, School
of Geology, Aristotle University of Thessaloniki, Greece, Special Volume 102:101.
Lazaridis, G., and E. Tsoukala. 2015. Comparative anatomy of carpal and tarsal bones in
three sympatric Turolian hipparions (Kryopigi, Chalkidiki, Greece). Bollettino della
Società Paleontologica Italiana, 54.
Legendre, S. 1986. Analysis of mammalian communities from the late Eocene and Oligocene
of southern France. Palaeovertebrata, 16(4):191-212.
Legendre, S. 1989. Les communautés de mammifères du Paléogène (Eocène supérieur et
Oligocène) d’Europe occidentale: structures, milieux et évolution. Münchener
Geowissenschaftliche abhandlungen, Reihe A, Geologie und Paläontologie, 16:1-110.
Lehmann, U. 1950. Über Mastodontenreste in der Bayerischen Staatssammlung in München.
Palaeontographica Abteilung A:121-228.
Lewis, G. E. 1939. A new Bramatherium skull. American Journal of Science, 237(4):275-
Liu, L., D. S. Kostopoulos, and M. Fortelius. 2004. Late Miocene Microstonyx remains
(Suidae, Mammalia) from Northern China. Geobios, 37(1):49-64.
Liu, L., D. S. Kostopoulos, and M. Fortelius. 2005. Suidae (Mammalia, Artiodactyla) from
the late Miocene of Akkaşdağı, Turkey. Geodiversitas, 27(4):715-733.
Liu, T., C. Li, and R. Zhai. 1978. Pliocene vertebrates of Lantian, Shensi. Professional Papers
of Stratigraphy and Palaeontology, 7:149-200.
Lopatin, A. V., A. S. Tesakov, and V. V. Titov. 2003. Late Miocene – early Pliocene
porcupines (Rodentia, Hystricidae) from South European Russia. Russian Journal of
Theriology, 2(1):26–32.
Lydekker, R. 1877. Notices of new and rare Mammals from the Siwaliks. Records of the
Geological Survey of India, 10:76-83.
Lydekker, R. 1878. Notices of Siwalik mammals. Records of the Geological Survey of India,
(1):64-104.
Lyman, R. L. 2008. Quantitative Paleozoology. Cambridge University Press.
Lyras, G. A. 2009. The evolution of the brain in Canidae (Mammalia: Carnivora). Scripta
Geologica, 139:1-94.
Lyras, G. A., and A. A. Van Der Geer. 2003. External brain anatomy in relation to the
phylogeny of Caninae (Carnivora: Canidae). Zoological Journal of the Linnean
Society, 138(4):505-522.
MacLeod, N., and K. D. Rose. 1993. Inferring locomotor behavior in Paleogene mammals via
Eigenshape analysis. American Journal of Science, 293(A):300-355.
Mandic, O., M. Harzhauser, K. F. Kuiper, D. Nagel, J. R. Wijbrans, E. Tsoukala, and G.
Lazaridis. in press. History of the Paratethys-Mediterranean gateway - age constrains
from mollusk stratigraphy and 40Ar/39Ar geochronology (N Greece). Geology.
Markov, G. N. 2004. The fossil proboscideans of Bulgaria and the importance of some
Bulgarian finds–a brief review. Historia naturalis bulgarica, 16:139-150.
Markov, G. N. 2008. The Turolian proboscideans (Mammalia) of Europe: preliminary
observations. Historia naturalis bulgarica, 19:153-178.
Martin, R. E. 1999. Taphonomy: a process approach. Cambridge University Press, 4.
Melentis, J. K. 1967. Die Pikermi Fauna von Halmyropotamos (Euböa, Griechenland). I Teil:
Odontologie und Kraniologie. Annales Géologiques des Pays Helleniques, 19:283-
Melentis, J. K. 1969. Die Pikermifauna von Halmyropotamos (Euböa, Griechenland). II Teil:
Osteologie. Annales Géologiques des Pays Helleniques, 21:217-306.
Melentis, J. K. 1974. Helladotherium duvernoyi Gaudry, 1860 aus Pikerml (Griechenland).
Sci. Annals, Fac. Phys. & Mathern., Univ. 'I'hessaloniki, 14:65-73.
Melentis, J. K., and H. Schneider. 1966. Studien über Fossile Vertebraten Griechenlands. 17.
Eine neue Pikermifauna in der Nähe der Ortschaft Alifaka in Thessalien
(Griechenland). Annales Géologiques des Pays Helleniques, 17:267-288.
Merceron, G., D. S. Kostopoulos, L. de Bonis, F. Fourel, G. D. Koufos, C. Lecuyer, and F.
Martineau. 2013. Stable isotope ecology of Miocene bovids from Northern Greece and
the ape/monkey turnover in the Balkans. J Hum Evol, 65(2):185-198.
Merceron, G., G. D. Koufos, and X. Valentin. 2009a. Feeding habits of the first European
colobine, Mesopithecus (Mammalia, Primates): evidence from a comparative dental
microwear analysis with modern cercopithecids. Geodiversitas, 31(4):865-878.
Merceron, G., J. Scott, R. S. Scott, D. Geraads, N. Spassov, and P. S. Ungar. 2009b. Folivory
or fruit/seed predation for Mesopithecus, an earliest colobine from the late Miocene of
Eurasia. J Hum Evol, 57(6):732-738.
Michaux, J. 1969. Muridae (Rodentia) du Pliocene superieur d'Espagne et du Midi de la
France. Palaeovertebrata, 3:1-25.
Mlikovsky, J. 1996. Tertiary Avian Localities of Greece. Tertiary avian localities of Europe.-
Acta universitatis Carolinae Geologica, 39:649-653.
Montoya, P. 1993. The porcupine Hystrix suevica Schlosser, 1884 from the lower Turolian of
Crevillente 2 (Spain). Scripta Geol, 103:135-149.
Morlo, M. 1999. Niche structure and evolutionin creodont (Mammalia) faunas of the
European and North American Eocene. Geobios, 32(2):297-305.
Morlo, M., G. Gunnell, and D. Nagel. 2010. Ecomorphological analysis of carnivore guilds in
the Eocene through Miocene of Laurasia. New contributions to the natural history of
Carnivora:269-310.
Morlo, M., and D. Nagel. 2007. The carnivore guild of the Taatsiin Gol area: Hyaenodontidae
(Creodonta), Carnivora, and Didymoconida from the Oligocene of Central Mongolia.
Annalen des Naturhistorischen Museums in Wien. Serie A für Mineralogie und
Petrographie, Geologie und Paläontologie, Anthropologie und Prähistorie, 108A:217-
Mottl, M. 1966. Neue Säugetierfunde aus dem Jungtertiär der Steiermark, VII: ein
vollständiger Hyotherium palaeochorus-Schädel aus dem Altpliozän (Pannon Südost-
Oesterreichs, p. 73-102.Volume 28. Mitteilungen des Museum für Bergbau Geologie
und Technik am Landesmuseum "Joanneum" Graz.
Nagel, D., and G. D. Koufos. 2009. The Late Miocene Mammal Faunas of the Mytilinii
Basin, Samos Island, Greece: New Collection. 15. Carnivore Guild Structure. Beiträge
zur Paläeontologie, 31:391–396.
Nagel, D., and M. Morlo. 2003. Guild structure of the carnivorous mammals (Creodonta,
Carnivora) from the Taatsiin Gol area, Lower Oligocene of Central Mongolia.
Deinsea, 10:419-429.
Osborn, H. F. 1936. Proboscidea. A monograph of the discovery, evolution, migration and
extinction of the mastodonts and elephants of the world, Volume I: Moeritherioidea,
Deinotherioidea, Mastodontoidea. American Museum Press, New York.
Owen, R. 1848. Description of teeth and portions of jaws of two extinct anthracotherioid
quadrupeds (Hyopotamus vectianus and Hyop. bovinus) discovered by the
Marchioness of Hastings in the Eocene deposits on the NW coast of the Isle of Wight:
with an attempt to develop Cuvier’s idea of the classification of pachyderms by the
number of their toes. Quarterly Journal of the Geological Society of London,
(1):104-141.
Ozansoy, F. 1965. Étude des gisements continentaux et des mammifères du Cénozoïque de
Turquie. Mémoires de la Société Géologique de France, 44:1-92.
Palmqvist, P., B. Martinez-Navarro, and A. Arribas. 1996. Prey selection by terrestrial
carnivores in a lower Pleistocene paleocommunity. Paleobiology, 22(4):514-534.
Pavlow, M. 1908. Quelques carnivores fossiles du gouvernement de Kherson et de
Bessarabie. Memoires de la Societe des Natumlistes de la Nouoeue-Russie, Odessa,
:27-48.
Pavlow, M. 1913. Mammifères tertiaires de la nouvelle Russie. Nouv. Mém. Soc. d. Natural.
Moscou, 17.
Pavlow, M. 1914. Aperçu sur la nuvelle faune des mammiferes tertiaires de la Russie
méridionale. Annuaire géologique et mineralogique de la Russie, 16:181-191.
Pérez-García, A., and E. Vlachos. 2014. New generic proposal for the European Neogene
large testudinids (Cryptodira) and the first phylogenetic hypothesis for the medium
and large representatives of the European Cenozoic record. Zoological Journal of the
Linnean Society, 172(3):653-719.
Pesquero, M. D., L. Alcalá, and Y. Fernández‐Jalvo. 2013. Taphonomy of the reference
Miocene vertebrate mammal site of Cerro de la Garita, Spain. Lethaia, 46(3):378-398.
Pickford, M. 1988. Revision of the Miocene Suidae of the Indian subcontinent. Münchner
Geowissenschaftliche abhandlungen, Reihe A, Geologie und Paläontologie, 12:1-91.
Pickford, M. 2013. Conohyus simorrensis (Lartet, 1851) (Suidae, Mammalia) from the
Middle Miocene of Carpetana (Madrid, Spain). Spanish Journal of Palaeontology,
(1):91-102.
Pickford, M. 2015. Late Miocene Suidae from Eurasia: the Hippopotamodon and Microstonyx
problem revisited Münchener Geowissenschaftliche abhandlungen, Reihe A, Geologie
und Paläontologie, 42:1-126.
Pickford, M., and L. Liping. 2001. Revision of the Miocene Suidae of Xiaolongtan (Kaiyuan),
China. Bolletino della Società Paleontologica Italiana, 40(2):275-284.
Pilgrim, G. E. 1910. Notices of new mammalian genera and species from the Tertiaries of
India. Records of the Geological Survey of India, 40(1):63-71.
Pilgrim, G. E. 1926a. The fossil suidae of India. Memoirs of the Geological Survey of India,
:1-65.
Pilgrim, G. E. 1926b. On the names and types of certain Pontian antelopes. Annals and
Magazine of Natural History, s. 9, 18:464.
Pilgrim, G. E. 1931. Catalogue of the Pontian Carnivora of Europe in the Department of
Geology. British Museum (Natural History), London, 174 p.
Pilgrim, G. E. 1933. A fossil skunk from Samos. American Museum novitates, 663(1-15).
Pilgrim, G. E. 1937. Siwalik Antelopes and Oxen in the American Museum of Natural
History. Bulletin of the American Museum of Natural History, 72(7):729-891.
Pilgrim, G. E. 1939. The fossil Bovidae of India. Paleontologia Indica n.s., 1:1-365.
Pilgrim, G. E., and A. T. Hopwood. 1928. Catalogue of the Pontian Bovidae of Europe in the
Department of Geology. British Museum (Natural History), London.
Plummer, T. W., and L. C. Bishop. 1994. Hominid paleoecology at Olduvai Gorge, Tanzania
as indicated by antelope remains. J Hum Evol, 27(1):47-75.
Qiu, Z.-X., B. Wang, and G. Xie. 2000. Preliminary report on a new genus of Ovibovinae
from Hezheng district, Gansu, China. Vertebrata Palasiatica, 38(2):131-140.
Radović, P., S. Alaburić, Z. Marković, and S. Vlastić. 2013. New view on the old
collection:'Pikermian fauna'from the vicinity of Veles (Republic of Macedonia), Part
: Primates. Bulletin of the Natural History Museum(6):7-29.
Rage, J.-C., and S. Sen. 1976. Les amphibiens et les reptiles du Pliocène supérieur de Çalta
(Turquie). Géologie Méditerranéenne, 3:127-134.
Rey, K., R. Amiot, C. Lécuyer, G. D. Koufos, F. Martineau, F. Fourel, D. S. Kostopoulos, and
G. Merceron. 2013. Late Miocene climatic and environmental variations in northern
Greece inferred from stable isotope compositions (δ18O, δ13C) of equid teeth apatite.
Palaeogeography, Palaeoclimatology, Palaeoecology, 388:48-57.
Rodler, A. 1888. Notiz über ein Besitze des Dr J.E. Polak beindliches Schädelfragment eines
Sivatheriden vom Knochenfelde von Maragha am Urmiasee in nordpersien. Anzeiger
Österreichische Akademia der Wissenschaften Mathematisch naturwissenschaftliche
Klasse, 25:114-115.
Rodler, A., and K. A. Weithofer. 1890. Die Wiederkäuer der Fauna von Maragha.
Denkschriften der Mathematisch - Naturwissenschaftlichen Classe der Kaiserlichen
Akademie Wissenschaften, 57:753-771.
Rögl, F., R. L. Bernor, M. D. Dermitzakis, C. Müller, and M. Stancheva. 1991. On the
Pontian correlation in the Aegean (Aegina Island). Newsletters on Stratigraphy:137-
Rook, L. 1999. Late Turolian Mesopithecus (Mammalia, Primates, Colobinae) from Italy. J
Hum Evol, 36(5):535-547.
Roth, J. R., and A. J. Wagner. 1854. Die fossilen Knochen-Ueberreste von Pikermi in
Griechenland. Abhandlungen der Bayerischen Akademie der Wissenschaften,
Mathematisch-Physikalische Klasse, 7:371-464.
Roussiakis, S. Metailurus major Zdansky, 1924 (Carnivora, Mammalia) from the classical
locality of Pikermi (Attica, Greece). Annales de Paléontologie, 87:119-132.
Roussiakis, S. 2002. Musteloids and feloids (Mammalia, Carnivora) from the late Miocene
locality of Pikermi (Attica, Greece). Geobios, 35(6):699-719.
Roussiakis, S. Prostrepsiceros and Protragelaphus (Artiodactyla, Mammalia) from the Late
Miocene locality of Chomateri (Attica, Greece). Annales de Paléontologie, 95:181-
Roussiakis, S., and G. Theodorou. 2001. Ancylotherium pentelicum (Gaudry & Lartet, 1856)
(Perissodactyla, Mammalia) from the classic locality of Pikermi (Attica, Greece),
stored in the Palaeontological and Geological Museum of Athens. Geobios, 34(5):563-
Roussiakis, S., and G. Theodorou. 2003. Carnivora from the Late Miocene of Kerassia
(Northern Euboea, Greece). Deinsea, 10:469-497.
Roussiakis, S., G. Theodorou, and G. Iliopoulos. 2006. An almost complete skeleton of
Metailurus parvulus (Carnivora, Felidae) from the late Miocene of Kerassia (Northern
Euboea, Greece). Geobios, 39(4):563-584.
Sakai, S. T., B. M. Arsznov, B. L. Lundrigan, and K. E. Holekamp. 2011. Virtual endocasts:
an application of computed tomography in the study of brain variation among hyenas.
Annals of the New York Academy of Sciences, 1225(S1):E160-E170.
Sanders, W. J. 2003. Proboscidea, p. 202-219 In M. Fortelius, J. Kappelman, S. Sen, and R.
L. Bernor (eds.), Geology and Paleontology of the Miocene Sinap Formation, Turkey.
Columbia University Press, New York.
Schaub, S. 1943. Die Vorderextremit at von Ancylotherium pentelicum Gaudry and Lartet.
Schweizerischen Paläontologischen Abhandlungen, 64:1-36.
Schlesinger, G. 1917. Die Mastodonten des K. K. Naturhistorischen Hofmuseums.
Denkschriften des K. K. Naturhistorischen Hofmuseums
, 230 p.
Schlesinger, G. 1922. Die Mastodonten der Budapester Sammlungen. Geologica Hungarica,
(1):1-284.
Schlosser, M. 1903. Die fossilen Säugethiere Chinas nebst einer Odontographie der recente
Antilopen. Abh. kön. bayr. Akad. Wiss., math.-physik. Kl., XXII, Abt, 1:1-221.
Schlosser, M. 1904. Die fossilen Cavicornia von Samos. Beiträge zur Paläeontologie und
Geologie Österreich-Ungrans, 17:21-118.
Schmidt-Kittler, N., H. De Bruijn, and C. Doukas. 1995. General introduction, p. 9-18. In N.
Schmidt-Kittler (ed.), The vertebrate locality of Maramena (Macedonia, Greece) at the
Turolian-Ruscinian boundary (Neogene).Volume 28. Münchner Geowissenschaftliche
Abhandlungen, München.
Scott, R. S. 2004. The Comparative Paleoecology of Late Miocene Eurasian Hominoids,
University of Texas 1-458 p.
Scott, R. S., and M. Maga. 2005. Paleoecology of the Akkaşdağı hipparions (Mammalia,
Equidae), late Miocene of Turkey, p. 809-830. In S. Sen (ed.), Geology, mammals and
environments at Akkașdagı, late Miocene of Central Anatolia.Volume 27.
Geodiversitas
Sein, C., J. van der Made, and G. E. Rössner. 2009. New material of Propotamochoerus
(Suidae, Mammalia) from the Irrawaddy Formation, Myanmar. Neues Jahrbuch für
Geologie und Paläontologie-Abhandlungen, 251(1):17-31.
Semenov, Y. 1988. Ictitherium spelaeum sp. n. (Carnivora, Viverridae) du Sarmatien moyen
d' Ukraine. Vestnik Zoologii, 4:49-59.
Semenov, Y. 1989. Iktiterii i mofologicheski skhodnye hieny neogena SSSR. [Ictitheres and
morphologically related hyaenas from the Neogene of the USSR.] 178 pp. Naukova
Dumka, Kiev. Russian.
Semenov, Y. 2008. Taxonomical reappraisal of “ictitheres”(Mammalia, Carnivora) from the
Late Miocene of Kenya. Comptes Rendus Palevol, 7(8):529-539.
Sen, S. 1999. 42 Family Hystricida, p. 427-434. In G. E. Rössner and K. Heissig (eds.), The
Miocene Land Mammals of Europe. Verlag Dr. Friedrich Pfeil, München.
Sen, S. 2001a. Early Pliocene porcupine (Mammalia, Rodentia) from Perpignan, France: a
new systematic study. Geodiversitas, 23(2):303-312.
Sen, S. 2001b. Rodents and Insectivores from the Upper Miocene of Molayan, Afghanistan.
Palaeontology, 44(5):913-932.
Sen, S., G. Bouvrain, and D. Geraads. 1998. Pliocene vertebrate locality of Calta, Ankara,
Turkey. 12. Paleoecology, biogeography and biochronology. Geodiversitas,
(3):497-510.
Sen, S., L. de Bonis, N. Dalfes, D. Geraads, and G. Koufos. 1994. Les gisements de
mammifères du Miocène supérieur de Kemiklitepe, Turquie: 1. Stratigraphie et
magnétostratigraphie. Bull. Mus. Natl. Hist. Nat., Paris, 4eme Ser., Section C, 16:5-
Sen, S., G. D. Koufos, D. Kondopoulou, and L. de Bonis. 2000. Magnetostratigraphy of late
Miocene continental deposits of the Lower Axios valley, Macedonia, Greece.
Geological Society of Greece, Special Publications, 9:197-206.
Sen, S., and Z. Purabrishemi. 2010. First porcupine fossils (Mammalia, Rodentia) from the
late Miocene of NW Iran, with notes on late Miocene–Pliocene dispersal of
porcupines. Paläontologische Zeitschrift, 84(2):239-248.
Shipman, P. 1981. Life history of a fossil. An introduction to taphonomy and palaeoecology.
Harvard University Press, England.
Sickenberg, O. 1932. Eine neue Antilope, Parurmiatherium rugosifrons nov. gen. nov. sp.,
aus dem Unterpliozän von Samos. Anzeiger Osterreichische Akademia der
Wissenchaften Mathematisch naturwissenschaftliche Klasse, 1:10-11.
Simpson, G. G., and A. Roe. 1939. Quantitative zoology: Numerical concepts and methods in
the study of recent and fossil animals.
Snel, E., M. Mărunţeanu, and J. Meulenkamp. 2006. Calcareous nannofossil biostratigraphy
and magnetostratigraphy of the Upper Miocene and Lower Pliocene of the Northern
Aegean (Orphanic Gulf–Strimon Basin areas), Greece. Palaeogeography, Palaeoclimatology, Palaeoecology, 238(1):125-150.
Solounias, N. 1981a. The Turolian Fauna from the Island of Samos, Greece: With Special
Emphasis on the Hyaenids and the Bovids. S Karger Ag.
Solounias, N. 1981b. Τhe Turolian fauna from the island of Samos, Greece. Contribution to
vertebrate Evolution, 6:1-232.
Solounias, N., F. Rivals, and G. M. Semprebon. 2010. Dietary interpretation and
paleoecology of herbivores from Pikermi and Samos (late Miocene of Greece).
Journal Information, 36(1):113–136.
Sondaar, P. Y. The Samos Hipparion. Proceedings Koninklijke Nederlandse Akademie
Wetenschappen, Amsterdam u. London, B.74::417-441.
Spassov, N., and D. Geraads. 2004. Tragoportax PILGRIM, 1937 and Miotragocerus
STROMER, 1928 (Mammalia, Bovidae) from the Turolian of Hadjidimovo, Bulgaria,
and a revision of the late Miocene Mediterranean Boselaphini. Geodiversitas,
(2):339-370.
Spassov, N., and D. Geraads. 2014. A new felid from the Late Miocene of the Balkans and
the contents of the genus Metailurus Zdansky, 1924 (Carnivora, Felidae). Journal of
Mammalian Evolution:1-12.
Spassov, N., D. Geraads, L. Hristova, G. N. Markov, G. Merceron, T. Tzankov, K. Stoyanov,
M. Böhme, and A. Dimitrova. 2012. A hominid tooth from Bulgaria: the last pre-
human hominid of continental Europe. J Hum Evol, 62(1):138-145.
Steffens, P., H. De Bruijn, J. Meulenkamp, and L. Benda. 1979. Field guide to the Neogene of
Northern Greece (Thessaloniki area and Strimon basin). Publications of the
Department of Geology and Paleontology, University of Athens, Series A, 35:1-14.
Stehlin, H. G. 1899-1900. Über die Geschichte des Suiden-Gebisses. Abh. schweiz. palaont.
Ges., 26-27:1-528.
Ştiuca, E. 2003. Note préliminaire sur les mammifères du Miocène de Reghiu (Dept. Vrancea,
Roumanie). Advances in Vertebrate Paleontology" Hen to Panta:113-116.
Stoykova, K., I. Zagorchev, and J.-P. Suc. 2012. Siviris Formation (Upper Oligocene-Lower
Miocene), a new lithostratigraphic unit, Kassandras Peninsula (Chalkidiki, Northern
Greece). Comptes Rendus de l'Academie Bulgare des Sciences, 65(8):1117-1124.
Stromer, E. 1928. Wirbeltiere im obermiozänen Flinz Μünchens. Abhandlungen der
Bayerischen Akademie der Wissenschaften, Mathematisch-Physikalische Klasse,
:1-71.
Studer, T. 1911. Eine neue Equidenform aus dem Obermiocän von Samos. Verhandlungen
der Deutschen Zoologischen Gesellschaft, 20-21:192-200.
Sulimski, A. 1960. Hystrix primigenia Wagner in the Pliocene fauna from Węże. Acta
Palaeontologica Polonica, 5(3):319-333.
Sylvestrou, I. A., and D. S. Kostopoulos. 2006. The late Miocene vertebrate locality of
Perivolaki, Thessaly, Greece. 7. Suidae. Palaeontographica Abteilung A:121-133.
Sylvestrou, I. A., and D. S. Kostopoulos. 2009. The late Miocene Mammal faunas of the
Mytilinii basin, Samos Island, Greece: new collection. 12. Suidae. Beitrage zur
Paläontologie, 31:283-297.
Symeonidis, N. 1973. Chalicotherium goldfussi Kaup (Perissodactyla, Mammalia) aus dem
Altpliozän von Pikermi (Griechenland). Annales Géologiques des Pays Helleniques,
:301-307.
Symeonidis, N. 1978. Ein Schädel von Metailurus parvulus (Hensel) aus Pikermi (Attica,
Griechenland). Annales Géologiques des Pays Helleniques, 29(2):698-703.
Tassy, P. 1983. Les Elephantoidea miocènes du plateau du Potwar, groupe de Siwalik,
Pakistan. Masson.
Tassy, P. 1994. Les gisements de mammifères du Miocène supérieur de Kemiklitepe,
Turquie. VII: Proboscidea (Mammalia). Bulletin du Muséum national d'histoire
naturelle. Section C, Sciences de la terre, paléontologie, géologie, minéralogie,
(1):143-157.
Tassy, P. 1996. Dental homologies and nomenclature in the Proboscidea. The Proboscidea:
Evolution and Palaeoecology of Elephants and Their Relatives. Oxford University
Press, Oxford:21-25.
Tassy, P. 2005. Proboscideans (Mammalia) from the late Miocene of Akkaşdağı.
Geodiversitas, 27(4):707-714.
Tassy, P., S. Sen, J.-J. Jaeger, J.-M. Mazin, and N. Dalfes. 1989. Une sous-espèce nouvelle de
Choerolophodon pentelici (Proboscidea, Mammalia) à Esme Akcaköy, Miocène
superiéur d'Anatolie occidentale. Comptes rendus de l'Académie des sciences. Série 2,
Mécanique, Physique, Chimie, Sciences de l'univers, Sciences de la Terre,
(20):2143-2146.
Thenius, E. 1950. Postpotamochoerus nov. subgen. hyotherioides aus dem Unterpliozän von
Samos (Griechenland) und die Herkunft der Potamochoeren. Sitzungsberichten der
Österreichischen Akademie der Wissenschaften Mathematisch-Naturwissenschaftliche
Klasse, 159:25-36.
Thenius, E. 1972. Microstonyx antiquus aus dem Alt-Pliozän Mittel-Europas. Zur Taxonomie
und Evolution der Suidae (Mammalia). Annalen des Naturhistorischen Museums in
Wien:539-586.
Theodorou, G. 1995. Observations on the carpus and tarsus of Hipparion from Pikermi
(Attica, Greece). Annales Géologiques des Pays Helleniques, 37:921-980.
Theodorou, G., A. Athanassiou, S. Roussiakis, and G. Iliopoulos. 2003. Preliminary remarks
on the late Miocene herbivores of Kerassia (Northern Euboea, Greece). Deinsea,
:519-530.
Theodorou, G., and S. Nicolaides. 1988. Stratigraphic horizons at the classic mammal locality
of Pikermi, Attica, Greece. Modern Geology, 13:177-181.
Theodorou, G., S. Roussiakis, A. Athanassiou, and A. Filippidi. 2010. Mammalian remains
from a new site near the classical locality of Pikermi (Attica, Greece). Scientific
Annals, School of Geology, Aristotle University of Thessaloniki, Sp, 99:09-119.
Thomas, H., S. Sen, and G. Ligabue. 1980. La faune Miocène de la formation Agha Jari du
Jebel Hamrin (Irak). Proceedings of the Koninklijke Nederlandse Akademie van
Wetenschappen, 83:269-287.
Tobien, H. 1973. On the evolution of mastodonts (Proboscidea, Mammalia). Hessisches
Landesamt für Bodenforschung.
Tobien, H. 1980. A note on the skull and mandible of a new choerolophodonte mastodont
(Probosc
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